New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam)

Jaap J. Vermeulen, e-mail:

JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands

Hong Truong Luu, e-mail:

Southern Institute of Ecology, Vietnam Academy of Science and Technology, 1 Mac Dinh Chi, Ho Chi Minh, Vietnam

Keum Theary, e-mail:

Cambodian Entomology Initiatives, Royal University of Phnom Penh, 12101 Phnom Penh, Cambodia

Katja Anker

JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands

Twenty-two new species of land snails are described from limestone hills on the west flank of the Mekong river delta, in Cambodia and Vietnam. The species belong to the families Assimineidae, Cyclophoridae, Diplommatinidae, Ariophantidae, Helicarionidae, Streptaxidae and Vertiginidae.

Key words
land snails; Cambodia; Vietnam; Mekong Delta Limestone Hills

Crosse H. 1863. Descriptions d’espèces nouvelles de Poulon Condor (Cochinchine). Journal de Conchyliologie, Paris 11: 269–273.
Greķe K. 2017. Taxonomic review of Diplommatinidae (Caenogastropoda: Cyclophoroidea) from Wallacea and the Papuan Region. In: Telnov D., Barclay M. V. L., Pauwels O. S. G. (eds). Biodiversity, biogeography and nature conservation in Wallacea and New Guinea. Volume 3. The Entomological Society of Latvia, Riga, pp. 151–316.
Dautzenberg P. 1894. Mollusques nouveaux recueillis au Tonkin par M. le capitaine Em. Dorr. Journal de Conchyliologie, Paris 41: 157–165.
Dautzenberg P., Fischer P. H. 1905. Liste des Mollusques récoltés par M. Mansuy en Indo-Chine et au Yunnan et description d’espèces nouvelles, II. Journal de Conchyliologie, Paris 53: 343–471.
Godwin Austen H. H. 1897–1914. Land and freshwater Mollusca of India, 2. Porter, London.
IUCN Red List – The IUCN (International Union of Conservation of Nature) Red List of Threatened Species, available at
Liew T. S., Vermeulen J. J., Marzuki M. E., Schilthuizen M. 2014. A cybertaxonomic revision of the micro-­landsnail genus Plectostoma Adam (Mollusca, Caeno­gastropoda, Diplommatinidae), from Peninsular Mala­ysia, Sumatra and Indochina. ZooKeys 393: 1–107.
Maassen W. J. M. 2001. Four new Diplommatinidae (Gastropoda, Prosobranchia, Diplommatinidae) from southern Thailand and northern Peninsular Malaysia. Basteria 65: 51–56.
Martens E. von 1908. Beschreibung einiger im östlichen Borneo von Dr. Martin Schmidt gesammelten Land- und Süsswasser-Conchylien. Mitteilungen aus dem Zoologischen Museum in Berlin 4: 252–291.
Möllendorff O. F. von 1885. Materialien zur Fauna von China. Die Auriculaceen, Nachträge und Berichtungen. Jahrbücher der Deutschen Malakolozoologischen Gesellschaft 12: 349–398.
Möllendorff O. F. von 1897. Neue Landschnecken von Java. Nachrichtsblatt der Deutschen Malakolo­zoologischen Gesellschaft 29: 57–72.
Montfort P. D. de 1810. Conchyliologie systématique, et classification méthodique des coquilles. Coquilles univalves, non cloissonneés, 2. F. Schoell, Paris.
Panha S., Burch J. B. 2002. New pupilloid land snails from Thailand (Pulmonata: Pupillidae). Natural History Journal Chulalongkorn University 2: 21–24.
Philippi R. A. 1846. Vier neue Konchylienarten. Zeitschrift für Malakozoologie 1846: 191–192.
Rensch B. 1931. Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. Zoologisches Jahrbüch Abteilung für Systematik, Ökologie und Geographie der Tiere 61: 361–396.
Swainson W. 1840. A treatise on malacology; or the natural classification of shells and shellfish. Lardner’s Cabinet Cyclopedia, London.
Tanmuangpak K., Tumpeesuwan C., Tumpeesuwan S. 2017. A new species of Sesara Albers, 1860 from northeastern Thailand (Stylommatophora: Helicarionidae). Molluscan Research 37: 1–7.
Van Benthem Jutting W. S. S. 1952. The Malayan species of Opisthostoma (Gastropoda, Prosobranchia, Cyclophoridae), with a catalogue of all the species hitherto described. Bulletin of the Raffles Museum, Singapore 24: 5–62.
Van Benthem Jutting W. S. S. 1958. Landmollusken von Sumba. Verhandlungen der Naturforschenden Gesellschaft, Basel 69: 90–117.
Van Benthem Jutting W. S. S. 1962. Coquilles terrestres nouvelles de quelques calcaires du Cambodge et du Sud Vietnam. Journal de Conchyliologie, Paris 102: 3–15.
Vermeulen J. J., Hong Truong Luu, Keum Theary, Anker K. in press. Revision of Notharinia (Mollusca: Gastropoda Caenogastropoda Diplommatinidae) of the Mekong Delta Limestone hills (Cambodia, Vietnam). Bulletin of the Raffles Museum, Singapore.
Vermeulen J. J., Liew T. S., Schilthuizen M. 2015. Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species. ZooKeys 531: 1–139.
Vermeulen J. J., Phung C. L., Truong Q. T. 2007. New species of terrestrial molluscs (Caenogastropoda, Pupinidae & Pulmonata: Vertiginidae) of the Hon Chong – Ha Tien limestone hills, Southern Vietnam. Basteria 71: 81–92.
Vermeulen J. J., Whitten A. J. 1998. Fauna Malesiana – Guide to the land snails of Bali. Backhuys Publishers & Fauna Malesiana Foundation, Leiden.

Folia Malacologica (2019) 27: 7-41
First published on-line: 2019-02-11 00:00:00
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